Maximum extent of Indo-Greek territory circa 175 BC (Source).
The Indo-Greek Kingdom (or sometimes Greco-Indian Kingdom) covered various parts of the northwest and northern Indian subcontinent from 180 BC to around 10 AD, and was ruled by a succession of more than thirty Greek kings, often in conflict with each other. The kingdom was founded when the Greco-Bactrian king Demetrius invaded India in 180 BC, ultimately creating an entity which seceded from the powerful Greco-Bactrian Kingdom centered in Bactria (today's northern Afghanistan). The city of Taxila in northern Pakistan would eventually serve as the Kingdom's cosmopolitan capital and seat of power.
During the two centuries of their rule, the Indo-Greek kings combined the Greek and Indian languages and symbols, as seen on their coins, and blended Ancient Greek, Hindu and Buddhist religious practices, as seen in the archaeological remains of their cities and in the indications of their support of Buddhism. The Indo-Greek kings seem to have achieved a level of cultural syncretism with no equivalent in history, the consequences of which are still felt today, particularly through the diffusion and influence of Greco-Buddhist art.
The Indo-Greeks ultimately disappeared as a political entity around 10 AD following the invasions of the Indo-Scythian, Indo-Parthian and Kushans, although pockets of Greek populations probably remained for several centuries longer.
The founder of the Indo-Greek Kingdom Demetrius I (205-171 BC), wearing the scalp of an elephant, symbol of his conquest of India.
Main article:History of the Indo-Greek Kingdom
Local turmoil preceded the invasion of northern India undertaken by Demetrius, son of the Greco-Bactrian king Euthydemus, circa 180 BC. General Pusyamitra Sunga had destroyed the ruling Mauryan King and had recently founded the Sunga dynasty (18578 BC). Written evidence of the initial Greek invasion survives in the Greek writings of Strabo and Justin, and in Sanskrit in the records of Patanjali, Kālidāsa, and in the Yuga Purana, among others. Coins and architectural evidence also attest to the extent of the initial Greek campaign.
Evidence of the initial invasion
According to Strabo, Greek advances temporarily went as far as the Sunga capital Pataliputra (today Patna) in eastern India:
"Those who, after Alexander, advanced beyond the Hypanis, to the Ganges and Pataliputra." (Strabo, 15-1-27) 
The 1st century BC Greek historian Apollodorus, quoted by Strabo, affirms that the Bactrian Greeks, led by Demetrius I and Menander, conquered India and occupied a larger territory than the Macedonians under Alexander the Great, going beyond the Hypanis towards the Himalayas.
The Roman historian Justin also mentionned the Indo-Greek conquests, describing Demetrius as "King of the Indians" ("Regis Indorum"), and explaining that after vanquishing him Eucratides in turn "put India under his rule" ("Indiam in potestatem redegit"). Although "India" only meant the upper Indus for Alexander the Great, since the embassies of Megasthenes in the 3rd century BC "India" meant to the Greeks most of the northern half of the Indian subcontinent, an area roughly corresponding to the extent of the Mauryan Empire at its largest.
To the south, the Greeks occupied the areas of the Sindh and Gujarat down to the region of Surat (Greek: Saraostus) near Mumbai (Bombay), including the strategic harbour of Barigaza (Bharuch), as attested by several writers (Strabo 11; Periplus of the Erythraean Sea, Chap. 41/47) and as evidenced by coins dating from the Indo-Greek ruler Apollodotus I:
"The Greeks... took possession, not only of Patalena, but also, on the rest of the coast, of what is called the kingdom of Saraostus and Sigerdis." (Strabo 11.11.1)
The 1st century AD Periplus of the Erythraean Sea describes numerous Greek buildings and fortifications in Barigaza, although mistakenly attributing them to Alexander, and the circulation of Indo-Greek coinage in the region:
"In these places there remain even to the present time signs of the expedition of Alexander, such as ancient shrines, walls of forts and great wells." Periplus, Chap. 41
"To the present day ancient drachmae are current in Barygaza, coming from this country, bearing inscriptions in Greek letters, and the devices of those who reigned after Alexander, Apollodorus and Menander." Periplus Chap. 47 
Various Indian records describe Greek (called Yavana) attacks on Mathura, Panchala, Saketa, and Pataliputra. Patanjali, a grammarian and commentator on Panini around 150 BC, describes in the Mahābhāsya, the invasion in two examples using the imperfect tense of Sanskrit, denoting a recent event:
"Arunad Yavanah Sāketam" ("The Yavana [Greeks] were besieging Saketa")
"Arunad Yavano Madhyamikām" ("The Yavana were besieging Madhyamika (the "Middle country").
The Anushasanaparava of the Mahabharata affirms that the country of Mathura, the heartland of India, was under the joint control of the Yavanas and the Kambojas. Accounts of battles between the Greeks and the Sunga in Central India are also found in the Mālavikāgnimitra, a play by Kālidāsa which describes a battle between Greek forces and Vasumitra, the grandson of Pushyamitra, during the latter's reign.
Also the Brahmanical text of the Yuga Purana, which describes Indian historical events in the form of a prophecy, relates the attack of the Indo-Greeks on the capital Pataliputra, a magnificent fortified city with 570 towers and 64 gates according to Megasthenes, and describes the ultimate destruction of the city's walls:
"Then, after having approached Saketa together with the Panchalas and the Mathuras, the Yavanas, valiant in battle, will reach Kusumadhvaja ("The town of the flower-standard", Pataliputra). Then, once Puspapura (another name of Pataliputra) has been reached and its celebrated mud[-walls] cast down, all the realm will be in disorder." (Yuga Purana, Paragraph 47-48, 2002 edition.)
According to the Yuga Purana a situation of complete social disorder follows, in which the Yavanas rule and mingle with the people, and the position of the Brahmins and the Sudras is inverted:
"Sudras will also be utterers of bho (a form of address used towards an equal or inferior), and Brahmins will be utterers of arya (a form of address used towards a superior), and the elders, most fearful of dharma, will fearlessly exploit the people. And in the city the Yavanas, the princes, will make this people acquainted with them: but the Yavanas, infatuated by war, will not remain in Madhyadesa." (Yuga Purana, Paragraph 55-56, 2002 edition.)
Indo-Greek stone palette representing an Hellenistic Nereid goddess riding a Ketos sea-monster, 2nd century BC, Sirkap.
The city of Sirkap, today in northwestern Pakistan, was built according to the "Hippodamian" grid-plan characteristic of Greek cities, suggesting it may have been built by Demetrius. Numerous Hellenistic artifacts have been found, in particular coins of Greco-Bactrian kings and stone palettes representing Greek mythological scenes. Some of them are purely Hellenistic, others indicate an evolution of the Greco-Bactrian styles found at Ai-Khanoum towards more indianized styles. For example, accessories such as Indian ankle bracelets can be found on some representations of Greek mythological figures such as Artemis. Various Buddhist structures, such as the Butkara Stupa in the area of Swat were decorated with Hellenistic architectural elements from the 2nd century BC.
Ancient fortifications have been unearthed in Rajagriha, south of Pataliputra, which are similar in structure to the Indo-Greek fortifications of Sirkap and Barikot: stone walls connected by jutting rectangular towers, reinforced by a low structure to repel siege machinery, in contrast to the mud walls used throughout the country at that time.
Tetradrachm of Menander I in Greco-Bactrian style (Alexandria-Kapisa mint) [Source].
Indian-standard coin of Apollodotus I (180-160 BC).
The first invasion was completed by 175 BC, as the Indo-Greeks contained the Sungas to the area eastward of Pataliputra, and established their rule on the new territory. Back in Bactria however, around 170 BC, an usurper named Eucratides managed to topple the Euthydemid dynasty. He took for himself the title of king and started a civil war by invading the Indo-Greek territory, forcing the Indo-Greeks to retreat from their easternmost possessions and establish their new oriental frontier at Mathura, to confront this new threat:
"The Yavanas, infatuated by war, will not remain in Madhadesa (the Middle Country). There will be mutual agreement among them to leave, due to a terrible and very dreadful war having broken out in their own realm." (Yuga Purana, paragraphs 56-57, 2002 edition).
The Hathigumpha inscription, written by the king of Kalinga, Kharavela, also describes the presence of the Greek king "Demetrius" with his army in eastern India, apparently as far as the city of Rajagriha about 70 km southeast of Pataliputra and one of the foremost Buddhist sacred cities, but claims that Demetrius ultimately retreated to Mathura on hearing of Kharavela's military successes further south:
"Then in the eighth year, (Kharavela) with a large army having sacked Goradhagiri causes pressure on Rajagaha (Rajagriha). On account of the loud report of this act of valour, the Yavana (Greek) King Dimi[ta] retreated to Mathura having extricated his demoralized army and transport." Hathigumpha inscription, in Epigraphia Indica, Vol. XX.
In any case, Eucratides seems to have occupied territory as far as the Indus, between ca 170 BC and 150 BC. His advances were ultimately checked by the Indo-Greek king Menander I (Milinda), previously a general of Demetrius, who asserted himself in the Indian part of the empire, apparently conquered Bactria as indicated by his issue of coins in the Greco-Bactrian style, and even began the last expansions eastwards.
Consolidation and rise of Menander I
Menander (Milinda) is considered as probably the most successful Indo-Greek king, and the conqueror of the vastest territory. The finds of his coins are the most numerous and the most widespread of all the Indo-Greek kings. In Antiquity, from at least the 1st century AD, the "Menander Mons", or "Mountains of Menander", came to designate the mountain chain at the extreme east of the Indian subcontinent, today's Naga hills and Arakan, as indicated in the Ptolemy world map of the 1st century AD geographer Ptolemy. Presumably the "Menander Mons" were so named because they marked the easternmost extent of his Indian conquests. Menander is also remembered in Buddhist literature (the Milinda Panha) as a convert to Buddhism: he became an arhat whose relics were enshrined in a manner reminiscent of the Buddha. He also introduced a new coin type, with Athena Alkidemos ("Protector of the people") on the reverse, which was adopted by most of his successors in the East.
Conquests east of the Punjab region were most likely made during the second half of the century by the king Menander I.
Following Menander's reign, about twenty Indo-Greek kings are known to have ruled in succession in the eastern parts of the Indo-Greek territory. Upon his death, Menander was succeeded by his queen Agathokleia, who for some time acted as regent to their son Strato I.
Greek provinces or satrapies in India
From ancient authors (Pliny, Arrian, Ptolemy and Strabo), a list of ten provinces, satrapies, or simple regional designations from within the Indo-Greek Kingdom can be discerned (though others have been lost):
From 130 BC, the Scythians and then the Yuezhi, following a long migration from the border of China), started to invade Bactria from the north. Around 125 BC the Greco-Bactrian king Heliocles, son of Eucratides, was probably killed during the invasion and the Greco-Bactrian kingdom proper ceased to exist. Heliocles may have been survived by his relative Eucratides II, who ruled south of the Hindu Kush, in areas untouched by the invasion. Other Indo-Greek kings like Zoilos I, Lysias and Antialcidas may possible have been relatives of either the Eucratid or the Euthydemid dynasties; they struck both Greek and bilingual coins and established a kingdom of their own.
A stabilizing alliance with the Yuezhi then seems to have followed, as hinted on the coins of Zoilos I, who minted coins showing Heracles' club together with a steppe-type recurve bow inside a victory wreath.
The Indo-Greeks thus suffered encroachments by the Greco-Bactrians in their western territories. The Indo-Greek territory was divided into two realms: the house of Menander retreated to their territories east of the Jhelum River as far as Mathura, whereas the Western kings ruled a larger kingdom of Paropamisadae, western Punjab and Arachosia to the south.
Silver drachm of Menander I (160-135 BC).
Obv: Greek legend, BASILEOS SOTHROS MENANDROY lit. "Saviour King Menander".
Rev: Kharosthi legend: MAHARAJA TRATASA MENADRASA "Saviour King Menander". Athena advancing right, with thunderbolt and shield.
Buddhism flourished under the Indo-Greek kings, and it has been suggested, although direct evidence is lacking, that their invasion of India was intended to show their support for the Mauryan empire, which had a long history of marital alliances, treaties of friendship, and exchange of ambassadors and religious emissaries with the Greeks,  and to protect the Buddhist faith from the religious persecutions of the Sungas. The city of Sirkap founded by Demetrius combines Greek and Indian influences without signs of segregation between the two cultures.
The first Greek coins to be minted in India, those of Menander I and Appolodotus I bear the mention "Saviour king" (BASILEOS SOTHROS), a title with high value in the Greek world which indicated an important deflective victory. For instance, Ptolemy I had been Soter (saviour) because he had helped save Rhodes from Demetrius the Besieger, and Antiochus I because he had saved Asia Minor from the Gauls.
Also, most of the coins of the Greek kings in India were bilingual, written in Greek on the front and in Pali (in the Kharoshthi script) on the back, a tremendous concession to another culture never before made in the Hellenic world. From the reign of Apollodotus II, around 80 BC, Kharoshthi letters started to be used as mintmarks on coins in combination with Greek monograms and mintmarks, suggesting the participation of local technicians to the minting process. Incidentally, these bilingual coins of the Indo-Greeks were the key in the decipherment of the Kharoshthi script by James Prinsep (1799-1840). Kharoshthi became extinct around the 3rd century AD.
In Indian literature, the Indo-Greeks are described as Yavanas (transliteration of "Ionians"). Direct epigraphical evidence involves the Indo-Greek kings, such as the mention of the "Yavana king" Antialcidas on the Heliodorus pillar in Vidisha, or the mention of Menander I in the Buddhist text of the Milinda Panha. In the Harivamsa the "Yavana" Indo-Greeks are qualified, together with the Sakas, Kambojas, Pahlavas and Paradas as Kshatriya-pungava i.e foremost among the Warrior caste, or Kshatriyas. The Majjhima Nikaya explains that in the lands of the Yavanas and Kambojas, in contrast with the numerous Indian castes, there were only two classes of people, Aryas and Dasas (masters and slaves). The Arya could become Dasa and vice versa.
In addition to the worship of the Classical pantheon of the Greek deities found on their coins (Zeus, Herakles, Athena, Apollo...), the Indo-Greeks were involved with local faiths, particularly with Buddhism, but also with Hinduism and Zoroastrianism.
Main article: Greco-Buddhism
The Edicts of Ashoka, inscribed during the reign of the Indian emperor Ashoka (273-232 BC), claim that the Greek populations of the northwestern Indian subcontinent (in today's Afghanistan and ancient Gandhara) had already welcomed Buddhism by around 250 BC:
"Here in the king's domain among the Greeks, the Kambojas, the Nabhakas, the Nabhapamkits, the Bhojas, the Pitinikas, the Andhras and the Palidas, everywhere people are following Beloved-of-the-Gods' instructions in Dharma. (Edicts of Ashoka, 13th Rock Edict, S. Dhammika).
After the Greco-Bactrians militarily occupied parts of northern India from around 180 BC, numerous instances of interaction between Greeks and Buddhism are recorded.
The conversion of Menander
A Hellenistic-style Buddhist stupa in the Indo-Greek city of Sirkap, northern Pakistan, 2nd century BC. It combines the sculptures of three temple-fronts: Greek (hidden from view, on the left), Hindu (center) and Buddhist (right). (Wider view:  (http://www.web.virginia.edu/asianarc/public/taxila/eagle02.jpg)).
Menander I, one of the most famous successors of Demetrius, ruled from 150 to 135 BC. He is presented by Greek authors as an even greater conqueror than Alexander the Great. Strabo (XI.II.I) says Menander was one of the two Bactrian kings who extended their power farthest into India.
Menander, the "Saviour king", seems to have converted to Buddhism, and is described in Buddhist texts as a great benefactor of the religion, on a par with Ashoka or the future Kushan emperor Kanishka. He is famous for his dialogues with the Buddhist monk Nagasena, transmitted to us in the Milinda Panha, which explain that he became a Buddhist arhat:
"And afterwards, taking delight in the wisdom of the Elder, he handed over his kingdom to his son, and abandoning the household life for the house-less state, grew great in insight, and himself attained to Arahatship!" (The Questions of King Milinda, Translation by T. W. Rhys Davids)
Plutarch also presents Menander as an example of benevolent rule, and explains that upon his death, the honour of sharing his remains was claimed by the various cities under his rule, and they were enshrined in "monuments" (probably stupas), in a parallel with the historic Buddha:
"But when one Menander, who had reigned graciously over the Bactrians, died afterwards in the camp, the cities indeed by common consent celebrated his funerals; but coming to a contest about his relics, they were difficultly at last brought to this agreement, that his ashes being distributed, everyone should carry away an equal share, and they should all erect monuments to him." (Plutarch, "Political Precepts" Praec. reip. ger. 28, 6).
During the reign of Menander, the Greek (Pali: Yona, lit: "Ionian") Buddhist monk Mahadhammarakkhita (Sanskrit: Mahadharmaraksita, lit. "Great protector of the Dharma") is said to have come from Alasandra (thought to be Alexandria of the Caucasus, the city founded by Alexander the Great, near today's Kabul) with 30,000 monks for the foundation ceremony of the Maha Thupa ("Great stupa") built by king Dutthagamani at Anuradhapura in Sri Lanka, indicating the importance of Buddhism within Greek communities in northwestern India, and the prominent role Greek Buddhist monks played in them:
"From Alasanda the city of the Yonas came the thera (elder) Yona Mahadhammarakkhita with thirty thousand bhikkhus." (Mahavamsa, XXIX)
Several Buddhist dedications by Greeks in India are recorded, such as that of the Greek meridarch (civil governor of a province) named Theodorus, describing in Kharoshthi how he enshrined relics of the Buddha. The inscriptions were found on a vase inside a stupa, dated to the reign of Menander or one his successors in the 1st century BC (Tarn, p391):
"Theudorena meridarkhena pratithavida ime sarira sakamunisa bhagavato bahu-jana-stitiye":
"The meridarch Theodorus has enshrined relics of Lord Shakyamuni, for the welfare of the mass of the people"
(Swāt relic vase inscription of the Meridarkh Theodoros)
Although the spread of Buddhism to Central Asia and Northern Asia is usually associated with the Kushans, a century or two later, there is a possibility that it may have been introduced in those areas from Gandhara "even earlier, during the time of Demetrius and Menander" (Puri, "Buddhism in Central Asia").
From around 180 BC, Agathocles and Pantaleon, probable successors to Demetrius I in the Paropamisadae, and the earliest Greek kings to issue Indian-standard square bilingual coins (in Brahmi), depicted the Buddhist lion together with the Hindu goddess Lakshmi.
Some coins of Agathocles in the Indian standard also show a six-arched stupa and a tree in a railing, typical symbol of the Bodhi tree in early Buddhism. These coins show an unprecedented willingness to adapt to every aspect of the local culture: shape of the coinage, coinage size, language, and religion.
Later, some Indo-Greek coins incorporate the Buddhist symbol of the eight-spoked wheel, such as those of Menander I, as well as his probable grandson Menander II. On these coins, the wheel is associated with the Greek symbols of victory, either the palm of victory, or the victory wreath handed over by the goddess Nike.
The ubiquitous symbol of the elephant may or may not have been associated with Buddhism. Interestingly, on some coin series of Antialcidas, the elephant holds the same relationship to Zeus and Nike as the Buddhist wheel on the coin of Menander II, tending to suggest a common meaning for both symbols. Some of the earlier coins of king Apollodotus I directly associate the elephant with Buddhist symbolism, such as the stupa hill surmounted by a star, also seen, for example on the coins of the Mauryan Empire or those of the later Kuninda kingdom. Conversely, the bull is probably associated with Shiva, and often described in an erectile state as on the coins of Apollodotus I.
Also, after the reign of Menander I, several Indo-Greek rulers, such as Agathokleia, Amyntas, Nicias, Peukolaos, Hermaeus, Hippostratos and Menander II, depicted themselves or their Greek deities forming with the right hand a symbolic gesture identical to the Buddhist vitarka mudra (thumb and index joined together, with other fingers extended), which in Buddhism signifies the transmission of the Buddha's teaching.
At precisely the same time, right after the death of Menander, several Indo-Greek rulers also started to adopt on their coins the Pali title of "Dharmikasa", meaning "follower of the Dharma" (the title of the great Indian Buddhist king Ashoka was Dharmaraja "King of the Dharma"). This usage was adopted by Strato I, Zoilos I, Heliokles II, Theophilos, Peukolaos, Menander II and Archebios.
Altogether, the conversion of Menander I to Buddhism suggested by the Milinda Panha seems to have triggered the use of Buddhist symbolism in one form or another on the coinage of close to half of the kings who succeeded him. Especially, all the kings after Menander who are recorded to have ruled in Gandhara (apart from the little known Demetrius III) display Buddhist symbolism in one form or another. On the contrary, none of the kings whose rule was limited to Punjab did display Buddhist signs (with the exception of the powerful Hippostratos, who probably took under his protection many Gandharan Greeks fleeing from the Indo-Scythians (Tarn).).
A 2nd century BC relief from a Buddhist stupa in Bharhut, in eastern Madhya Pradesh (today at the Indian Museum in Calcutta), represents a foreign soldier with the curly hair of a Greek and the royal headband with flowing ends of a Greek king. In his left hand, he hold a branch of ivy, symbol of Dionysos. Also parts of his dress, with rows of geometrical folds, are characteristically Hellenistic in style. On his sword appears the Buddhist symbol of the three jewels, or Triratana.
Vitarka Mudra gestures on Indo-Greek coinage. Top: Divinities Tyche and Zeus. Bottom: Depiction of Indo-Greek kings Nicias and Menander II.
Representation of the Buddha
One of the first known representations of the Buddha, Gandhara, in pure Hellenistic style and technique.
The anthropomorphic representation of the Buddha is absent from Indo-Greek coinage, suggesting that the Indo-Greek kings may have respected the Indian aniconic rule for Buddhist depictions, limiting themselves to Buddhist symbolism only. Consistently with this perspective, the actual depiction of the Buddha would be a later phenomenon, usually dated to the 1st century AD, emerging from the sponsorship of the syncretic Kushan Empire and executed by Greek, and, later, Indian and possibly Roman artists. Datation of Greco-Buddhist statues is generally uncertain, but they are at least firmly established from the 1st century AD.
Another possibility is that the Indo-Greeks may not have considered the Buddha strictly as a God, but rather as an essentially human sage or philosopher, in line with the traditional Nikaya Buddhist doctrine. Just as philosophers were routinely represented in statues (but certainly not on coins) in Antiquity, the Indo-Greek may have initiated anthropomorphic representations of the Buddha in statuary only, possibly as soon as the 2nd-1st century BC, as advocated by Foucher and suggested by Chinese murals depicting Emperor Wu of Han worshipping Buddha statues brought from Central Asia in 120 BC (See picture). Foucher especially considered Hellenistic free-standing Buddhas as "the most beautiful, and probably the most ancient of the Buddhas", assigning them to the 1st century BC, and making them the starting point of the anthropomorphic representations of the Buddha.
The willingness of ancient Greeks to represent local deities is also attested in Egypt with the creation of the god Serapis in Hellenistic style, an adaptation of the Egyptian god Apis. An Indo-Chinese tradition also explains that Nagasena, also known as Menander's Buddhist teacher, created in 43 BC in the city of Pataliputra a statue of the Buddha, the Emerald Buddha, which was later brought to Thailand.
Stylistically, Indo-Greek coins generally display a very high level of Hellenistic artistic realism, which declined drastically around 50 BC with the invasions of the Indo-Scythians, Yuezhi and Indo-Parthians. The first known statues of the Buddha are also very realistic and Hellenistic in style and are more consistent with the pre-50 BC artistic level seen on coins. This would tend to suggest that the first statues were created between 130 BC (death of Menander) and 50 BC, precisely at the time when Buddhist symbolism appeared on Indo-Greek coinage. From that time, Menander and his successors may have been the key propagators of Buddhist ideas and representations: "the spread of Gandhari Buddhism may have been stimulated by Menander's royal patronage, as may have the development and spread of Gandharan sculpture, which seems to have accompanied it" (Mc Evilly, "The shape of ancient thought", p378)
The representation of the Buddha may also be connected to his progressive deification, which is usually associated with the spread of the Indian principle of Bhakti (personal devotion to a deity). Bhakti is a principle which evolved in the Bhagavata religious movement, and is said to have permeated Buddhism from about 100 BC, and to have been a contributing factor to the representation of the Buddha in human form. The association of the Indo-Greeks with the Bhagavata movement is documented in the inscription of the Heliodorus pillar, made during the reign of the Indo-Greek king Antialcidas (r.c. 115-95 BC). At that time relations with the Sungas seem to have improved, and some level of religious exchange seems to have occurred. The point of time when bhakti fervour would have encountered the Hellenistic artistic tradition would then be around 100 BC.
Most of the early images of the Buddha (especially those of the standing Buddha) are anepigraphic, which makes it difficult to have a definite datation. The earliest known image of the Buddha with approximate indications on date is the Bimaran casket, which has been found buried with coins of the Indo-Scythian king Azes II (or possibly Azes I), indicating a 30-10 BC date, although this date is not undisputed. Such datation, as well as the general Hellenistic style and attitude of the Buddha on the Bimaran casket (himation dress, contrapposto attitude, general depiction) would made it a possible Indo-Greek work, used in dedications by Indo-Scythians right after the end of Indo-Greek rule in the area of Gandhara. Since it already displays quite a sophisticated iconography (Brahma and Śakra as attendants, Bodhisattvas) in an advanced style, it would suggest much earlier representations of the Buddha were already current by that time, going back to the rule of the Indo-Greeks (Alfred A. Foucher and others).
The first known bilingual coins of the Indo-Greeks were issued by Agathocles around 180 BC. These coins were found in Ai-Khanoum, the great Greco-Bactrian city in northeastern Afghanistan, but introduce for the first time an Indian script (the Brahmi script which had been in use under the Mauryan empire), and the first known representations of Hindu deities, in a very Indian iconography: Krishna-Vasudeva, with his large wheel with six spokes (chakra) and conch (shanka), and his brother Samkarsa-Balarama, with his plough (hala) and pestle (masala), both early avatars of Vishnu. The square coins, instead of the usual Greek round coins, also followed the Indian standard for coinage. The dancing girls on some of the coins of Agathocles and Pantaleon are also sometimes considered as representations of Subhadra, Krishna's sister.
These first issues were in several respects a short-lived experiment. Hindu anthropomorphic deities were never again represented in Indo-Greek coinage (although the bull on the vast quantity of subsequent coins may have symbolized Shiva, as the elephant may have symbolized Buddhism), and the Brahmi script was immediately replaced by the Kharoshti script, derived from Aramaic. The general practice however of minting bilingual coins and combining Greek and Indian iconography, sometimes in the Greek and sometimes in the Indian standard continued for the next two centuries.
In any case, these coins suggest the strong presence of Indian religious traditions in the northwestern Indian subcontinent at that time, and the willingness of the Greeks to acknowledge and even promote them. Artistically, they tend to indicate that the Greeks were not particularly reluctant to make representations of local deities, which has some bearing on the later emergence of the image of the Buddha in Hellenistic style.
The Heliodorus pillar inscription is another epigraphical evidence of the interaction between Greeks and Hinduism. The pillar was erected around 110 BC in central India at the site of Vidisha, by Heliodorus, a Greek ambassador of the Indo-Greek king Antialcidas to the court of the Sunga king Bhagabhadra. The pillar was surmounted by a sculpture of Garuda and was apparently dedicated by Heliodorus to the temple of Vasudeva.
"This Garuda-standard of Vasudeva (Vishnu), the God of Gods
was erected here by the Bhagavata Heliodoros,
the son of Dion, a man of Taxila,
sent by the Great Greek (Yona) King
Antialkidas, as ambassador to
King Kasiputra Bhagabhadra, the Savior
son of the princess from Benares, in the fourteenth year of his reign."
(Heliodorus pillar inscription)
Persian culture and religion seem to have been rather influential among the Western Indo-Greeks, who, located around the Paropamisadae, lived in direct contact with the Central Asian cultural sphere and the eastern reaches of the Parthian empire. Images of the Persian Zoroastrian god Mithra appear extensively on the Indo-Greek coinage of the Western kings, as a god with a radiated phrygian cap.
This Zeus-Mithra god is also the one represented seated (with the rays around the head, and a small protrusion on the top of the head representing the cap) on many coins of Hermaeus, Antialcidas or Heliokles II, or possibly even earlier during the time of Eucratides I, on whose coins the deity is said to be the god of the city of Kapisa.
The future Buddha Maitreya, usually represented seated on a throne Western-style, and venerated both in Mahayana and non-Mahayana Buddhism, is sometimes considered as influenced by the god Mithra. "Some scholars suggest he (Maitreya) was originally linked to the Iranian saviour-figure Mitra, and that his later importance for Buddhist as the future Buddha residing in the Tusita heaven, who will follow on from Sakyamuni Buddha, derives from this source." (Keown, Dictionary of Buddhism)
Incipient Greco-Buddhist art
Main article: Greco-Buddhist art
The Buddha with Herakles/Vajrapani (left detail) and Tyche/Hariti (right detail) may be "incipient Buddhist sculpture in Indo-Greek style" (Boardman). Herakles still has his lion skin on the left shoulder, although his club has been replaced by Vajrapani's thunderbolt. Tyche holds a Classical cornucopia, Tapa-i-Shotor, Hadda, eastern Afghanistan (Image Source).
In general, the art of the Indo-Greeks is poorly documented, and few works of art (apart from their coins and a few stone palettes) are directly attributed to them. Traditionally, no sculptural remains have been attributed to the Indo-Greeks, although their Hellenistic heritage and artistic proficiency would naturally have encouraged such creations (as neighbouring and contemporary Ai-Khanoum abundantly suggests). On the contrary, and rather paradoxically, most Gandharan Hellenistic works of art are usually attributed to the direct successors of the Indo-Greeks in India, such as the Indo-Scythians, the Indo-Parthians and the Kushans, from the 1st century AD.
The possibility of a direct connection between the Indo-Greeks and Greco-Buddhist art has been reaffirmed recently as the dating of the rule of Indo-Greek kings has been extended to the first decades of the 1st century AD, with the reign of Strato II in the Punjab. Also, Foucher, Tarn and more recently Boardman or McEvilley have taken the view that some of the most purely Hellenistic works of northwestern India and Afghanistan, may actually be wrongly attributed to later centuries, and instead belong to a period one or two centuries earlier, to the time of the Indo-Greeks in the 2nd-1st century BC.  This is particularly the case of some purely Hellenistic works in Hadda, Afghanistan, an area which "might indeed be the cradle of incipient Buddhist sculpture in Indo-Greek style". Referring to one of the Buddha triads in Hadda (drawing), in which the Buddha is sided by very Classical depictions of Herakles/Vajrapani and Tyche/Hariti, Boardman explains that both figures "might at first (and even second) glance, pass as, say, from Asia Minor or Syria of the first or second century BC (...) these are essentially Greek figures, executed by artists fully conversant with far more than the externals of the Classical style". Many of the works of art at Hadda can also be compared to the style of the 2nd century BC sculptures of the Hellenistic world, such as those of the Temple of Olympia at Bassae in Greece, which could also suggest roughly contemporary dates.
Alternatively, it has been suggested that these works of art may have been executed by itinerant Greek artists during the time of maritime contacts with the West from the 1st to the 3rd century AD.
The supposition that such highly Hellenistic and, at the same time Buddhist, works of art belong to the Indo-Greek period would be consistent with the known Buddhist activity of the Indo-Greeks (the Milinda Panha etc...), their Hellenistic cultural heritage which would naturally have induced them to produce extensive statuary, their know artistic proficiency as seen on their coins until around 50 BC, and the dated appearance of already complex iconography incorporating Hellenistic sculptural codes with the Bimaran casket in the early 1st century AD.
Indo-Greeks in the art of Gandhara
Hellenistic culture in the Indian subcontinent: Greek clothes, amphoras, wine and music (Detail of Chakhil-i-Ghoundi stupa, Hadda, Gandhara, 1st century AD.
The Greco-Buddhist art of Gandhara, beyond the omnipresence of Greek style and stylistic elements which might be simply considered as an enduring artistic tradition, offers numerous depictions of people in Greek Classical realistic style, attitudes and fashion (clothes such as the chiton and the himation, similar in form and style to the 2nd century BC Greco-Bactrian statues of Ai-Khanoum, hairstyle), holding contraptions which are characteristic of Greek culture (amphoras, "kantaros" Greek drinking cups), in situations which can range from festive (such as Bacchanalian scenes) to Buddhist-devotional.
Uncertainties in dating make it unclear whether these works of art actually depict Greeks of the period of Indo-Greek rule up to the 1st century BC, or remaining Greek communities under the rule of the Indo-Parthians or Kushans in the 1st and 2nd century AD.
Very little is known about the economy of the Indo-Greeks. The abundance of their coins would tend to suggest large mining operations, particularly in the mountainous area of the Hindu-Kush, and an important monetary economy. The Indo-Greek did strike bilingual coins both in the Greek "round" standard and in the Indian "square" standard, suggesting that monetary circulation extended to all parts of society. The adoption of Indo-Greek monetary conventions by neighbouring kingdoms, such as the Kunindas to the east and the Satavahanas to the south, would also suggest that Indo-Greek coins were used extensively for cross-border trade.
It would also seem that some of the coins emitted by the Indo-Greek kings, particularly those in the monolingual Attic standard, may have been used to pay some form of tribute to the Yuezhi tribes north of the Hindu-Kush. This is indicated by the coins finds of the Qunduz hoard in northern Afghanistan, which have yielded quantities of Indo-Greek coins in the Hellenistic standard (Greek weights, Greek language), although none of the kings represented in the hoard are known to have ruled so far north. Conversely, none of these coins have ever been found south of the Hindu-Kush.
Trade with China
An indirect testimony by the Chinese explorer Zhang Qian, who visited Bactria around 128 BC, suggests that intense trade with Southern China was going through northern India, and therefore probably through the contemporary Indo-Greek realm. Zhang Qian explains that he found Chinese products in the Bactrian markets, and that they were transiting through India:
"When I was in Bactria," Zhang Qian reported, "I saw bamboo canes from Qiong and cloth (silk?) made in the province of Shu. When I asked the people how they had gotten such articles, they replied: "Our merchants go buy them in the markets of Shendu (northwestern India). Shendu, they told me, lies several thousand li southeast of Bactria. The people cultivate land, and live much like the people of Bactria". (Sima Qian, "Records of the Great Historian", trans. Burton Watson, p236).
Indian Ocean trade
Maritime relations across the Indian ocean started in the 3rd century BC, and further developed during the time of the Indo-Greeks together with their territorial expansion along the western coast of India. The first contacts started when the Ptolemies constructed the Red Sea ports of Myos Hormos and Berenike, with destination the Indus delta and the Kathiawar peninsula. Around 130 BC, Eudoxus of Cyzicus is reported (Strabo, Geog. II.3.4)  to have made a successful voyage to India and returned with a cargo of perfumes and gemstones. By the time Indo-Greek rule was ending, up to 120 ships were setting sail every year from Myos Hormos to India (Strabo Geog. II.5.12).
The coins of the Indo-Greeks provide rich clues on their uniforms and weapons. Typical Hellenistic uniforms are depicted, with helmets being either round in the Greco-Bactrian style, or the flat kausia of the Macedonians (coins of Apollodotus I).
Their weapons were spears, swords, longbow (on the coins of Agathokleia) and arrows. Interestingly, around 130 BC the Central Asian recurve bow of the steppes with its gorytos box starts to appear for the first time on the coins of Zoilos I, suggesting strong interactions (and apparently an alliance) with nomadic peoples, either Yuezhi or Scythian. The recurve bow becomes a standard feature of Indo-Greek horsemen by 90 BC, as seen on some of the coins of Hermaeus.
Generally, Indo-Greek kings are often represented riding horses, as soon as the reign of Antimachus II around 160 BC. The equestrian tradition probably goes back to the Greco-Bactrians, who are said by Polybius to have faced a Seleucid invasion in 210 BC with 10,000 horsemen. Although war elephants are never represented on coins, a harness plate (phalera) dated to the 3-2nd century BC, today in the Hermitage Museum, depicts a helmetted Greek combatant on an Indian war elephant, and would be either Greco-Bactrian or Indo-Greek work. Indian war elephants were a standard feature of Hellenistic armies, and this would naturally have been the case for the Indo-Greeks as well.
The Milinda Panha, in the questions of Nagasena to king Menander, provides a rare glimpse of the military methods of the period:
"(Nagasena) Has it ever happened to you, O king, that rival kings rose up against you as enemies and opponents?
-(Menander) Yes, certainly.
-Then you set to work, I suppose, to have moats dug, and ramparts thrown up, and watch towers erected, and strongholds built, and stores of food collected?
-Not at all. All that had been prepared beforehand.
-Or you had yourself trained in the management of war elephants, and in horsemanship, and in the use of the war chariot, and in archery and fencing?
-Not at all. I had learnt all that before.
-With the object of warding off future danger."
(Milinda Panha, Book III, Chap 7)
Size of Indo-Greek armies
The armed forces of the Indo-Greeks during their invasion of India must have been quite considerable, as suggested by their ability to topple local rulers, but also by the size of the armed reaction of some Indian rulers. The ruler of Kalinga, Kharavela, claims in the Hathigumpha inscription that he led a "large army" in the direction of Demetrius' own "army" and "transports", and that he induced him to retreat from Pataliputra to Mathura. A "large army" for the state of Kalinga must indeed have been quite considerable. The Greek ambassador Megasthenes took special note of the military strength of Kalinga in his Indica in the middle of the 3rd century BC:
"The royal city of the Calingae (Kalinga) is called Parthalis. Over their king 60,000 foot-soldiers, 1,000 horsemen, 700 elephants keep watch and ward in "procinct of war." (Megasthenes fragm. LVI. in Plin. Hist. Nat. VI. 21. 8-23. 11.). 
That this kind of military strength was needed to confront the Indo-Greeks is indicative of the Indo-Greeks' own military commitment.
An account by the Roman writer Justin gives another hint of the size of Indo-Greek armies, which, in the case of the conflict between the Greco-Bactrian Eucratides and the Indo-Greek Demetrius II, he numbers at 60,000 (although they allegedly lost to 300 Greco-Bactrians):
"Eucratides led many wars with great courage, and, while weakened by them, was put under siege by Demetrius, king of the Indians. He made numerous sorties, and managed to vanquish 60,000 enemies with 300 soldiers, and thus liberated after four months, he put India under his rule" (Justin, XLI,6)
The military strength of nomadic tribes from Central Asia (Yuezhi and Scythians) probably constituted a significant threat to the Indo-Greeks. According to Zhang Qian, the Yuezhi represented a considerable force of between 100,000 and 200,000 mounted archer warriors, with customs identical to those of the Xiongnu.
Finally, the Indo-Greek seem to have combined forces with other "invaders" during their expansion into India, since they are often referred to in combination with others (especially the Kambojas), in the Indian accounts of their invasions.
Eastern Indo-Greek territories
After around 100 BC, Indian kings recovered the area of Mathura and Eastern Punjab east of the Ravi River, and started to mint their own coins. The Arjunayanas (area of Mathura) and Yaudheyas mention military victories on their coins ("Victory of the Arjunayanas", "Victory of the Yaudheyas"). During the 1st century BC, the Trigartas, Audumbaras and finally the Kunindas (closest to Punjab) also started to mint their own coins, usually in a style highly reminiscent of Indo-Greek coinage.
The Western king Philoxenus briefly occupied the whole remaining Greek territory from the Paropamisadae to Western Punjab between 100 to 95 BC, after what the territories fragmented again. The eastern kings regained their territory as far west as Arachosia.
Around 80 BC, an Indo-Scythian king named Maues, possibly a general in the service of the Indo-Greeks, ruled for a few years in northwestern India before the Indo-Greeks again took control. King Hippostratos (65-55 BC) seems to have been one of the most successful subsequent Indo-Greek kings until he lost to the Indo-Scythian Azes I, who established an Indo-Scythian dynasty.
Throughout the 1st century BC, the Indo-Greeks progressively lost ground against the invasion of the Indo-Scythians. Although the Indo-Scythians clearly ruled militarily and politically, they remained surprisingly respectful of Greek and Indian cultures. Their coins were minted in Greek mints, continued using proper Greek and Kharoshthi legends, and incorporated depictions of Greek deities, particularly Zeus. The Mathura lion capital inscription attests that they adopted the Buddhist faith, as do the depictions of deities forming the vitarka mudra on their coins. Greek communities, far from being exterminated, probably persisted under Indo-Scythian rule.
The Indo-Greeks continued to rule a territory in the eastern Punjab, until the kingdom of the last Indo-Greek king Strato II was taken over by the Indo-Scythian ruler Rajuvula around 10 AD.
Western Indo-Greek territories
Around eight western Indo-Greek kings are known. The last important king was Hermaeus, who reigned until around 70 BC; soon after his death the Yuezhi took over his areas from neighbouring Bactria. Chinese chronicles (the Hou Hanshu) actually tend to suggest that the Chinese general Wen-Chung had helped negotiate the alliance of Hermaeus with the Yuezhi, against the Indo-Scythians. When Hermaeus is depicted on his coins riding a horse, he is equipped with the recurve bow and bow-case of the steppes.
After 70 BC, the Yuezhi became the new rulers of the Paropamisadae, and minted vast quantities of posthumous issues of Hermaeus up to around 40 AD, when they blend with the coinage of the Kushan king Kujula Kadphises. The first documented Yuezhi prince, Sapadbizes, ruled around 20 BC, and minted in Greek and in the same style as the western Indo-Greek kings, probably depending on Greek mints and celators.
Kushan man in traditional costume with tunic and boots, 2nd century AD, Gandhara.
The Yuezhi expanded to the east during the 1st century AD, to found the Kushan Empire. The first Kushan emperor Kujula Kadphises ostensibly associated himself with Hermaeus on his coins, suggesting that he may have been one of his descendants by alliance, or at least wanted to claim his legacy. The Yuezhi (future Kushans) were in many ways the cultural and political heirs to the Indo-Greeks, as suggested by their adoption of the Greek culture (writing system, Greco-Buddhist art) and their claim to a lineage with the last western Indo-Greek king Hermaeus.
The last known mention of an Indo-Greek ruler is suggested by an inscription on a signet ring of the 1st century AD in the name of a king Theodamas, from the Bajaur area of Gandhara, in modern Pakistan. No coins of him are known, but the signet bears in kharoshthi script the inscription "Su Theodamasa", "Su" being explained as the Greek transliteration of the ubiquitous Kushan royal title "Shau" ("Shah", "King").
Enduring legacy of the Indo-Greek Kingdom
From the 1st century AD, the Greek communities of central Asia and northwestern India lived under the control of the Kushan branch of the Yuezhi, apart from a short-lived invasion of the Indo-Parthian Kingdom. The Kushans founded the Kushan Empire, which was to prosper for several centuries. In the south, the Greeks were under the rule of the Western Kshatrapas.
It is unclear how much longer the Greeks managed to maintain a distinct presence in the Indian sub-continent.
At the beginning of the 2nd century AD, the Central India Satavahana king Gautamiputra (r. 106130 AD) adopted the epithet Sātakarni, meaning "Destroyer of Sakas (Western Kshatrapas), Yavanas (Indo-Greeks) and Pahlavas (Indo-Parthians)" in his inscriptions, suggesting a continued presence of the Indo-Greeks until that time.
Around 200 AD, the Manu Smriti describes the downfall of the Yavanas, as well as many others:
"43. But in consequence of the omission of the sacred rites, and of their not consulting Brahmanas, the following tribes of Kshatriyas have gradually sunk in this world to the condition of Shudras;
44. (Viz.) the Paundrakas, the Chodas, the Dravidas, the Kambojas, the Yavanas, the Shakas, the Paradas, the Pahlavas, the Chinas, the Kiratas, the Daradas and the Khashas." (Manusmritti, X.43-44)
The Brihat-Katha-Manjari text of the Sanskrit poet Kshmendra (11th and 12th centuries) (10/1/285-86) relates that around 400 AD the Gupta king Vikramaditya (Chandragupta II) had "unburdened the sacred earth of the Barbarians" like "the Shakas, Mlecchas, Kambojas, Yavanas, Tusharas, Parasikas, Hunas" etc… by annihilating these "sinners" completely.
A few common Greek words were adopted in Sanskrit, such as words related to writing and warfare:
"ink" (Sankrit: melā, Greek: μέλαν "melan")
"pen" (Sanskrit:kalamo, Greek:κάλαμος "kalamos")
"book" (Sanskrit: pustaka, Greek: πύξινον "puksinon")
a "horse's bit" (Sanskrit: khalina, Greek: χαλινός "khalinos")
a "siege mine" (used to undermine the wall of a fortress): (Sanskrit: surungā, Greek: σύριγγα "suringa")
The "Avaca" Kharosthi inscription, found on a Buddhist relic casket, indicates that the old Greek military title of strategos ("commander") had apparently endured the Indo-Scythian invasion and was being used by the Apracarajas of Bajaur during the 1st century AD (the inscription mentions the dedication date of the casket as "the year 63 of the late Maharaja Aya", Aya being the Indo-Scythian ruler Azes I, who started the Vikrama era in 58 BC, therefore suggesting a date around 5 AD). The dedication mentions "vaga stratego puyaite viyayamitro ya" i.e. "The Lord Commander (Stratego) Viyayamitra is honored too" 
The Greek philosopher Apollonius of Tyana is related by Philostratus in Life of Apollonius Tyana to have visited India, and specifically the city of Taxila around 46 AD. He describes constructions of the Greek type,  probably referring to Sirkap, and explains that the Indo-Parthian king of Taxila, named Phraotes, speaks Greek fluently, a language he had been educated in while in exile to the east, beyond the river Hyphasis:
"Tell me, O King, how you acquired such a command of the Greek tongue, and whence you derived all your philosophical attainments in this place?" 
[...]-"My father, after a Greek education, brought me to the sages at an age somewhat too early perhaps, for I was only twelve at the time, but they brought me up like their own son; for any that they admit knowing the Greek tongue they are especially fond of, because they consider that in virtue of the similarity of his disposition he already belongs to themselves." 
Lastly, from the inscription of Rabatak we have the following information, tending to indicate that Greek was still in official use until the time of Kanishka (circa 120 AD):
"He (Kanishka) issued(?) an edict(?) in Greek and then he put it into the Aryan language". …but when Kanishka refers to "the Aryan language" he surely means Bactrian, …"By the grace of Auramazda, I made another text in Aryan, which previously did not exist". It is difficult not to associate Kanishka's emphasis here on the use of the "Aryan language" with the replacement of Greek by Bactrian on his coinage. The numismatic evidence shows that this must have taken place very early in Kanishka's reign,…" Prof. Nicholas Sims-Williams (University of London).
The Greek language was probably still "alive" during the time of Kanishka, as this king introduced a new title in Greek on his early coins (BACIΛEΥC BACIΛEWΝ, "King of Kings" in the nominative), as well as the name of Greek deities such as HΛIOC (Sun god Helios), HΦAHCTOC (Fire god Hephaistos), and CAΛHNH (Moon goddess Selene).
The Greek script was used not only on coins, but also in manuscripts and stone inscriptions as late as the period of Islamic invasions in the 7th-8th century.
A Greek "Yona" calendar era seems to have been in use in Northwestern Indian for several centuries following the foundation of the Indo-Greek kingdom. A recently discovered inscription in Kharoshthi gives a relationship between several eras of the period:
"In the twenty-seventh - 27 - year in the reign of Lord Vijayamitra, the King of the Apaca; in the seventy-third - 73 - year which is called "of Azes", in the two hundred and first - 201 - year of the Yonas (Greeks), on the eighth day of the month of Sravana; on this day was established [this] stupa by Rukhana, the wife of the King of Apraca, [and] by Vijayamitra, the king of Apraca, [and] by Indravarma (Indravasu?), the commander (stratega), [together] with their wives and sons." (Senior 2003)
As the Azes era is usually considered as starting in 58 BC, the Yona era would correspond to 186 BC, which falls in the reign of Demetrius I, although dates ranging from 186 to 150 BC are still debated.
Astronomy and Astrology
One of the earliest Indian writings on astronomy and astrology (although not the earliest, as the Vedanga Jyotisha is dated to around 1350 BC), titled the Yavanajataka or "The Saying (Horoscopy to some) of the Greeks", is a translation from Greek to Sanskrit made by "Yavanesvara" ("Lord of the Greeks") in 149150 AD under the rule of the Western Kshatrapa king Rudrakarman I. The Yavanajataka contains instructions on calculating astrological charts (horoscopes) from the time and place of one's birth. Astrology flourished in the Hellenistic world (particularly Alexandria) and the Yavanajataka reflects astrological techniques developed in the Greek-speaking world. Astronomical mathematical methods, such as the calculate of the 'horoskopos' (zodiac sign on the eastern horizon) was used in the service of astrology.
Another set of treatises, the Paulisa Siddhanta and the Romaka Siddhantas, are attributed to later Greek influence in India. The Paulisa Siddhanta has been tentatively identified with the works of Paulus Alexandrinus, who wrote a well-known astrological hand-book.
Indian astronomy is widely acknowledged to be have been influenced by the Alexandrian school, and its technical nomenclature is essentially Greek: "The Yavanas are barbarians, yet the science of astronomy originated with them and for this they must be reverenced like gods" (The Gargi-Samhita). Several other Indian texts show appreciation for the scientific knowledge of the "Yavana" Greeks. 
Influence of Indo-Greek coinage
Overall, the coinage of the Indo-Greeks remained extremely influential for several centuries throughout the Indian subcontinent:
The Indo-Greek weight and size standard for silver drachms was adopted by the contemporary Buddhist kingdom of the Kunindas in Punjab, the first attempt by an Indian kingdom to produce coins that could compare with those of the Indo-Greeks.
In central India, the Satavahanas (2nd century BC- 2nd century AD) adopted the practice of representing their kings in profile, within circular legends.
The direct successors of the Indo-Greeks in the northwest, the Indo-Scythians and Indo-Parthians continued displaying their kings within a legend in Greek, and on the obverse Greek deities.
To the south, the Western Kshatrapas (1st-4th century AD) represented their kings in profile with circular legends in corrupted Greek.
The Kushans (1st-4th century AD) used the Greek language on their coinage until the first few years of the reign of Kanishka, whence they adopted the Bactrian language, written with the Greek script.
The Guptas (4th-6th century AD), in turn imitating the Western Kshatrapas, also showed their rulers in profile, within a legend in corrupted Greek, in the coinage of their western territories.
The latest use of the Greek script on coins corresponds to the rule of the Turkish Shahi of Kabul, around 850 AD. As late as the 13th century, the Sultan of Delhi Mohammad I (1295-1315), one of the first Muslim rulers of northern India, would use on his coins the title Sikander el-sani ("The second Alexander"), in a reference to his famous predecessor in the conquest of India.
Portraits from the site of Hadda, 3rd century AD.
Limited population genetics studies have been made on genetic markers such as mitochondrial DNA in the populations of the Indian subcontinent, to estimate the contribution of the Greeks to the genetic pool. Although some of the markers which are present in a large proportion of Greeks today have not been found, the Greek/European genetic contribution to the Punjab region has been estimated between 0%15%:
"The political influence of Seleucid and Bactrian dynastic Greeks over northwest India, for example, persisted for several centuries after the invasion of the army of Alexander the Great (Tarn 1951). However, we have not found, in Punjab or anywhere else in India, Y chromosomes with the M170 or M35 mutations that together account for 30% in Greeks and Macedonians today (Semino et al. 2000). Given the sample size of 325 Indian Y chromosomes examined, however, it can be said that the Greek homeland (or European, more generally, where these markers are spread) contribution has been 0%3% for the total population or 0%15% for Punjab in particular. Such broad estimates are preliminary, at best. It will take larger sample sizes, more populations, and increased molecular resolution to determine the likely modest impact of historic gene flows to India on its pre-existing large populations." (Kivisild et al. "Origins of Indian Casts and Tribes").
Some pockets of Greek populations probably remained for some time, and to this day, some communities in the Hindu Kush claim to be descendants of the Greeks, such as the Kalasha and Hunza in Pakistan, and the neighbouring Nuristani in Afghanistan.
One can not assume however that the present Greek population is representative of the Macedonian army under Alexander. This army probably contained a large number of Persians and other groups such as Scythians and Thracians.
Greco-Roman exchanges with India
Although the political power of the Greeks had waned in the north, mainly due to nomadic invasions, trade relations between the Mediterranean and India continued for several centuries. The trade started by Eudoxus of Cyzicus in 130 BC kept on increasing, and according to Strabo (II.5.12), by the time of Augustus, up to 120 ships were setting sail every year from Myos Hormos to India. So much gold was used for this trade, and apparently recycled by the Kushans for their own coinage, that Pliny (NH VI.101) complained about the drain of specie to India. In practice, this trade was still handled by Greek middlemen, as all the recorded names of ship captains for the period are Greek.
In India, the ports of Barbaricum (modern Karachi), Barygaza, and Muziris and Arikamedu on the southern tip of India were the main centers of this trade. Ptolemy described Muzuris as "a port packed with Greek ships" (VII.I.8), and the Tamil Sangam describes that "fine vessels, masterpieces of Yavana workmanship, arrive with gold and depart with pepper". In the south, the main trading partners were the Tamil dynasties of the Pandyas, Cholas and Cheras. The 2nd century Periplus of the Erythraean Sea describes Greco-Roman merchants selling in Barbaricum "thin clothing, figured linens, topaz, coral, storax, frankincense, vessels of glass, silver and gold plate, and a little wine" in exchange for "costus, bdellium, lycium, nard, turquoise, lapis lazuli, Seric skins, cotton cloth, silk yarn, and indigo". In Barygaza, they would buy wheat, rice, sesame oil, cotton and cloth. At Cape Comorin and in Sri Lanka, they would buy pearls and gems.
Also various exchanges are recorded between India and Rome during this period. In particular, embassies from India, as well as several missions from "Sramanas" to the Roman emperors are known (see Buddhism and the Roman world). Finally, Roman goods and works of art found their way to the Kushans, as archaeological finds in Begram have confirmed.
Main article:Silk Road transmission of Art
The "Kanishka casket", dated to the first year of Kanishka's reign in 127 AD, was signed by a Greek artist named Agesilas, who oversaw work at Kanishka's stupas (caitya), confirming the direct involvement of Greeks with Buddhist realizations at such a late date. (British Museum, drawing).
Greek representations and artistic styles, with some possible admixtures from the Roman world, continued to maintain a strong identity down to the 3rd4th century, as indicated by the archaeological remains of such sites as Hadda in eastern Afghanistan.
The Greco-Buddhist image of the Buddha was transmitted progressively through Central Asia and China until it reached Japan in the 6th century.
Numerous elements of Greek mythology and iconography, introduced in northwestern India by the Indo-Greeks through their coinage at the very least, were then adopted throughout Asia within a Buddhist context, especially along the Silk Road. The Japanese Buddhist deity Shukongoshin, one of the wrath-filled protector deities of Buddhist temples in Japan, is an interesting case of transmission of the image of the famous Greek god Herakles to the Far-East along the Silk Road. The image of Herakles was introduced in India with the coinage of Demetrius and several of his successors, used in Greco-Buddhist art to represent Vajrapani the protector of the Buddha, and was then used in Central Asia, China and Japan to depict the protector gods of Buddhist temples.
Another case of artistic transmission is the Greek Wind God Boreas, transiting through Central Asia and China to become the Japanese Shinto wind god Fujin. In consistency with Greek iconography for Boreas, the Japanese wind god holds above his head with his two hands a draping or "wind bag" in the same general attitude. The abundance of hair have been kept in the Japanese rendering, as well as exaggerated facial features.
Intellectual and religious legacy
Main article: Greco-Buddhism
The impact of the Indo-Greeks on Indian thought and religion is unknown, although many influences have been suggested. Scholars believe that Mahayana Buddhism as a distinct movement began around the 1st century BC in the North-western Indian subcontinent, corresponding to the time and place of Indo-Greek florescence. Intense multi-cultural influences have indeed been suggested in the appearance of Mahayana: "Key formative influences on the early development of the Mahayana and Pure Land movements, which became so much part of East Asian civilization, are to be sought in Buddhism's earlier encounters along the Silk Road". As Mahayana Buddhism emerged, it received "influences from popular Hindu devotional cults (bhakti), Persian and Greco-Roman theologies which filtered into India from the northwest". Many of the early Mahayana theories of reality and knowledge can be related to Greek philosophical schools of thought: Mahayana Buddhism has been described as "the form of Buddhism which (regardless of how Hinduized its later forms became) seems to have originated in the Greco-Buddhist communities of India, through a conflation of the Greek Democritean-Sophistic-Skeptical tradition with the rudimentary and unformalized empirical and skeptical elements already present in early Buddhism".
List of the Indo-Greek kings and their territories
Today 36 Indo-Greek kings are known. Several of them are also recorded in Western and Indian historical sources, but the majority are known through numismatic evidence only. The exact chronology and sequencing of their rule is still a matter of scholarly inquiry, with adjustments regular being made with new analysis and coin finds (overstrikes of one king over another's coins being the most critical element in establishing chronological sequences).
Lastly Theodamas, not mentionned in this list, may have been an Indo-Greek ruler in the Bajaur area in the 1st century AD.
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